◊ Aphodiinae
♦ Aphodiini
○ Aphodiina
♦ Psammodiini
○ Psammodiina
○ Rhyssemina
◊ Cetoniinae
♦ Cetoniini
○ Cetoniina
○ Leucocelina
◊ Dynastinae
♦ Oryctini
♦ Pentodontini
○ Pentodontina
◊ Hopliinae
♦ Hopliini
◊ Melolonthinae
♦ Melolonthini
○ Melolonthina
♦ Rhizotrogini
○ Rhizotrogina
◊ Rutelinae
♦ Adoretini
○ Adoretina
♦ Anomalini
○ Anisopliina
○ Anomalina
◊ Scarabaeinae
♦ Coprini
♦ Gymnopleurini
♦ Oniticellini
○ Oniticellina
♦ Onitini
○ Onitina
♦ Onthophagini
♦ Scarabaeini
♦ Sisyphini
◊ Sericinae
♦ Sericini
○ Sericina
◊ Trichiinae
♦ Trichiini
○ Trichiina
◊ Valginae
♦ Valgini
○ Valgina
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